For those who don't know,
this Wikipedia entry explains how the whole thing got started.
The basic nuts and bolts of it, is a typical otaku(shy, withdrawn, only able to express feelings online, etc.) was on a train one day, when some drunk guy stormed onto the car, and began harassing all the women on board. He decided to do something, and stood up to him, keeping him occupied until the conductor came, to take care of him. The women were all thankful for his efforts, but one in particular was deeply touched by his act, and asked if they could exchange addresses. He came to 2ch time and again for advice on what to do about the woman, how to treat her, what to do on their dates, and so on. Eventually, they confessed their love for each other, and this was cause for a mass celebration among the forums. Due to being a large influence on the Japanese culture, the story spread countrywide, and spawned anime, manga, books, and even a movie.